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Transcript
A
tropism (from the Greek trope — a turning) is growth or
turning in response to an environmental stimulus.
The ‘rhododendron thermometer’ (Figure 1) shows the response
of chilled leaves of rhododendrons to increased temperature. This
thermotropism — curling and drooping of the leaves when cold
— reduces water loss, and also reduces bleaching of chlorophyll
in bright sunlight.
Pollen grains show two tropisms. One is chemotropism —
growth in response to a chemical signal — they grow towards
the ovules, whatever the orientation of the flower. The second
is electrotropism — growth in response to an electrical field
(no-one knows why).
Heliotropism describes the movement of plant parts, such as
leaves and flowers, that track the sun as it goes across the sky.
Sunflower buds do this but once the flowers start to open they lose
this tropism, and end up all facing approximately east.
Thigmotropism is response to touch. When the extending
tendrils of a passion flower touch an object, they grow around it
so that the plant is secured and can climb higher.
Phototropism is one of the most familiar tropisms — shoots
grow towards the direction of light, but did you know that
plant seeds also show sonotropism? Beans germinate and grow
Dr Jeremy Burgess/SPL
Russ Drakeley
Tropisms
more quickly when exposed to ultrasound (probably due to the
ultrasound enhancing the activity of the enzyme alpha-amylase).
Gravi- or geotropism is also familiar — plant roots tend to
grow towards the direction of gravitational pull, but roots also
respond to gradients of humidity. It is not easy to detect this, as
soil is usually uniformly wet or dry, but experiments carried out
in microgravity, with plant roots grown in humid air, show that
root caps can detect gradients and signal roots to grow towards
water — hydrotropism.
Liz Sheffield
chemo- and electro9:20
10°C
helio-
9:10
–0.5°C
9:00
T = –3.0°C
Figure 1 The rhododendron thermometer.
Rhododendron on a warm spring day.
thigmo-
Rhododendron on a cold winter day.
gravi- and hydro-
Fotolia
thermo-
Biological Sciences Review
April 2013
Fotolia
Steve Gschmeissner/SPL
Lliz Sheffield
Fotolia
photo- and sono-