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Phylum Mollusca
BIO 2215
Oklahoma City Community College
Dennis Anderson
1
Phylum Mollusca
2
Numbers of species
Arthropoda
Mollusca
Chordata
Platyhelminthes
Nematoda
Annelida
Porifera
Echinodermata
Other
Sarcomastigophora
Apicomplex
Ciliophora
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Molluscs
• 50,000 -100,000 living
species
• 35,000 extinct species
• Largest = 1000 pounds
• 80% less than 5 cm
• Soft body
• Most have a shell
• Most marine
• Snails terrestrial
– Most habitat
4
Phylum Mollusca
• Ventral Foot
– Locomotion
• Mantle
– Encloses mantle cavity
• Shell
• Radula
• Coelom
(eucoelomate)
• Metanephridia
• Open circulatory
system
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– Closed in cephalopods
Economics
• Pearls
• Burrowing shipworms
• Snails & slugs
– Garden pests
– Food
– Intermediate hosts for
parasites
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Trochophore Larva
• Same type as Phylum
Annelida
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Uniramia
Echinodermata
Chelicerata
Vertebrata
Lophophores
Crustacea
Other Chordata
Arthropoda
Annelida
Hemichordata
Other
pseudocoelomates
Nematoda
Mesozoa
Sarcomastigophora
Ciliophora
Apicomplexa
Microspora
Mollusca
Nemertea
Platyhelminthes
Ctenophora
Cnidaria
Placozoa
Porifera
Myxozoa
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Generalized Mollusc
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Body Plan
Pericardial cavity
Metanephridium
Mantle cavity
Gonad
Ctenidium
Radula
Stomach and digestive gland
Foot
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Dorsal mantle covers the
visceral mass.
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Secretes the shell
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Ctenidium (Respiration)
13
Complete digestive system
14
Paired ventral nerve cords
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Radula
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Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display.
Radular Structure
Source: From A Life of Invertebrates, Copyright © 1979 W. D. Russell-Hunter.
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Coelom - metanephridia
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Class Polyplacophora
Chitons
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Class Polyplacophora
• Eight dorsal plates
• Reduced head
• Radula reinforced
with iron
– Scrape algae from
rocks
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Class Polyplacophora
Mouth
Mantle cavity
Ctenidium
Foot
Anus
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Class Polyplacophora
Digestive
gland
Mouth
Stomach
Gonad
Pericardial cavity
Nephridium
Anus
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Class Bivalvia
Clams, Oysters, Shipworms
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Class Bivalvia
•
•
•
•
Two shells
Most are filter feeders
No head or radula
Burrow
– Sand, wood, rocks
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Giant Clam & Burrowing Clam
Siphon
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Zebra Mussel
• Environmental Pest
• Ballast water of ships
from Europe in 1986
• Attack be secreting
adhesive byssal
threads
– Each other
– Other mussels
– Man made objects
• Pipes, plumbing
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Zebra Mussel
• Live in high densities
• Feed on
phytoplankton
• Reproduce rapidly
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Zebra Mussel
• Attach to native
mussels
• Killed all native
mussels in Lake Erie
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Distribution of Zebra Mussel
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Bivalve structures
Hinge
Labial palp
Ctenidium
Excurrent
siphon
Foot
Incurrent
siphon
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Clam anatomy
Pericardial cavity
Metanephridium
Heart
Intestine
Excurrent
Stomach
Incurrent
Intestine
Gonad
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Clam anatomy
Digestive
gland
Excurrent
Stomach
Incurrent
Intestine
Gonad
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Clam anatomy
Pericardial cavity
Heart
Intestine
Excurrent
Incurrent
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Clam anatomy
Metanephridium
Pericardial cavity
Heart
Intestine
Excurrent
Incurrent
Gonad
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Oysters
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Pearl formation
Shell
Developing pearl
Epithelium
Irritant lodged between shell and mantle
Layers of nacre secreted around foreign material
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Scallops
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Shipworms
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Class Gastropoda
Snails, Slugs, Conchs, Limpets
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Class Gastropoda
• One shell (if present)
• Torsion of body
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Snail
• Terrestrail
• Mantle cavity
functions as lung
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Snail
Shell
Pneumostome
Tentacle
(Eye stalks)
Tentacle
Anus
Foot
Mouth
Genital pore
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Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display.
Internal Structure of a Generalized Gastropod
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Nudibranch
• No shell
• Dorsal projections
– Gills
– Nematocyst discharge
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Abalone
• Several holes in top of shell
– Excrete waste
• Food for man
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Slug
• No shell
• Garden pests
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Limpet
• Herbivores
• Cling to rocks or other surfaces
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Conch
• Large shell
• Marine
• Many are predators
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Class Cephalopoda
Squids, Octopuses, Nautiluses
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Class Cephalopoda
• Shell in squid and
octopus absent or
vestigial
• Jet propulsion
• Ink sac
• Foot modified into
arms and tentacles
• Marine
• All predators
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Squid
Posterior surface
Right
Ventral
Dorsal
Left
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Squid
Tentacle
Arm
Funnel (siphon)
Collar
Eye
Fin
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Squid
Shell (Pen)
Systemic
heart
Branchial heart
Ctenidium
Funnel
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Squid Male
Testis
Penis
Hectocotylous arm
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Squid Female
Ovary with eggs
Oviducal gland
Nidamental glands
Oviducal opening
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Copyright © The McGraw-Hill Companies, Inc. Permission required for reproduction or display.
Cephalopod Eye
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Octopus
• Eight arms with
suckers
• Crawl or eject water
from siphon
• Change skin color
– chromatophores
• Most intelligent
invertebrate
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Nautilus
• Up to 94 tentacles
– No suckers
• Shell with many
chambers
– Lives in outermost
chamber
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Ammonoids
• Extinct
• Devonian to
Cretaceous
– 400 to 65 MYA
• Died out with
dinosaurs
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Class Scaphopoda
• Tooth shells
• Shell opens on both
ends
• Burrow into mud
• No gills
– Mantle for gas
exchange
• Feed on detritus and
protozoa
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Class Scaphododa
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The End
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