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Organismal Biology Organism-of-the-week: Pantherophis guttatus Common names: corn snake, red rat snake, chicken snake, etc. Arboreal=tree dwelling Species= Pantherophis guttatus o Common name: corn snake o Rules for species: Binomen= two name First name= Genus Always capitalized Always underlined OR italicized Plural: Genera Second name= Specific Epithet Not capitalized Always underlined OR italicized Genus= Pantherophis o Common name: rate snake o Plural: Genera Family= Colubridae o Non-venomous snakes o Family usually ends in –idae or –aceae Order= Squamata o All snakes and lizards o Suborder= serpentes Just snakes Suborders usually start with: Sub-, super-, Infra Class=Reptilia o Common name: reptiles Phlyum= Chordata o Common name: vertebrates and other chordates o Plural: Phyla Kingdom= Animalia o Common name: animals Biology= Study of life Bi= two o Plants and animals -ology= the study of Biodiversity= study of how organisms are different Survey Cladistics= common ancestry is the primary criterion used to classify organisms Clades= groups that contain an ancestral species and all of its descendents o Equal to a monophyletic group Monophyletic= single ancestor o Holophyletic= all descendents of the single ancestor o Paraphyletic= some but not all of descendents of the single ancestor o Monophylogeny is what we strive for Polyphyletic= more than one ancestor o (Not good) Plesiomorphic= around for a long time o Ancestral Apomorphic= newly evolved characteristic o Derived Symplesiomorphic= shared ancestral characteristic Synapomorphic= shared derived characteristics o This is what scientists want to avoid polyphyletic groups Autapomorphic= single unshared characteristic Systematics Taxonomy= the naming and classification of organisms o Taxon: the named taxonomic unit at any level of the hierarchy Example= snake Plural: taxa o Taxonomic revisions—why? (1) New information Molecular vs morphological Similarities among organisms o Analogous= common environment having characteristics that are similar because of convergent evolution Not ancestral traits Not used for classification o Homologous= come from same ancestors Similar in characteristics resulting from a shared ancestry Used for classification Ancestral traits What scientists want! (2)Different philosophy Dendogram (phylogram): family tree (3)Change: just because someone feels like changing it Phlogeny= evolutional history (natural selection) Classification= way of categorizing organisms Survey: 3 Domains, 6 Kingdoms, 35-40 Phyla Domains: Eukarya- True nucleus Archae- no nucleus or other membrane organelles Bacteria- no nucleus or other membrane organelles Eukarya: Kingdoms o Animalia Movement Cell membrane o Plantae Cell wall Autotrophic o o Fungi Cell wall Almost no movement Heterotrophs Protista Difficult to explain Variety Prokaryotes= Bacteria and Archae Not an official taxon The grouping of bacteria and archae as prokaryotes is incorrect because archaea is more related to eukarya than it is to bacteria Diversity o 2 domains o 2+ kingdoms Size o Small o Most are 1-5 m (micrometer) Shape o ⃝ –coccus Plural: cocci Spherical in shape o Oval shaped –bacillus Plural: bacilli o ~~-- Spirilla Plural: spirillae Solitary vs colonial Solitary= found alone Colonial= found in groups Streptoo OOOOOOO o One line Staphyloo Clusters Cell Wall o Present in most o Has a different chemistry than the one animals have o Variable Example: Gram Stain A staining method that distinguishes between two different kinds of bacterial cell wall o o Used to look at cell wall Positive/negative o Gram negative= describing the group of bacteria that have a cell wall that is structurally MORE complex and contains LESS peptidoglycan than the cell wall of a gram-positive bacteria. Gram negative bacteria are often more toxic than gram positive bacteria. o Gram positive= describing the group of bacteria that have a cell wall that is structurally LESS complex and contains MORE peptidoglycan than the cell wall of a gram-negative bacteria. Less toxic than gram-negative o Peptidoglycan is a type of polymer in bacterial cell walls consisting of modified sugars cross-linked by short polypeptides Capsule Fimbria Plural: fimbriae A short, hairlike appendage of a prokaryotic cell that helps it adhere to the substrate or to other cells o a substrate is the surface on which a plant or animal live Thinner and shorter than flagellum Pilus Plural: pili In bacteria, a structure that links one cell to another at the start of conjugation, also called a sex pilus or conjugation pilus o Conjugation is the direct transfer of DNA from one bacterial cell to another bacterial cell AKA “sex” pilus Motility o Flagellum A long cellular appendage specialized for locomotion There are different types Plural: flagella o Taxis= an oriented movement toward or away from a stimulus Phototaxis- move toward or away from light Chemotaxis- move toward or away from a chemical Genome o Nucleoid= a non-membrane enclosed region in a prokaryotic cell where its chromosome is located o Ring shaped DNA o Plasmids= a small circular double-stranded DNA molecule that carries accessory genes separate from those of a bacterial chromosome. Reproduction o Asexual reproduction No meiosis The generation of offspring from a single parent that occurs without the fusion of gametes. In most cases, the offspring are genetically identical to the parent o Binary fission Splits two ways A method of asexual reproduction by “division in half”. In prokaryotes binary fission does not involve mitosis (but it does in eukaryotes) Usually pretty rapid, but time varies o Budding o Endospores= a thick-coated, resistant cell produced by some bacterial cells when they are exposed to harsh conditions Very resistant Dormancy and dispersal Nutritional Mode o Heterotrophic= heterotrophs are organisms that obtain organic food molecules by eating other organisms or substances derived from them Decomposers= an organism that absorbs nutrients from nonliving organic material such as corpses, fallen plant material, and the wastes of living organisms and converts them to inorganic forms; aka a detritivore Pathogenic- causes disease o Autotrophic= autotrophs are organisms that obtain organic food molecules without eating other organisms or substances derived from other organisms. They use energy from the sun or from oxidation of inorganic substances to make organic molecules from inorganic ones. Photoautotrophic= capable of synthesizing their own food from inorganic substances (carbon dioxide) using light as an energy source Chemoautotrophic= an organism, typically a bacterium, that derives energy from the oxidation of inorganic compounds o Photoheterotrophs= organisms that use light for energy (ATP), but cannot use carbon dioxide as their sole carbon source. They must obtain carbon in organic form. Oxygen Relationships o Aerobic= with oxygen o Anaerobic= without oxygen o Facultative= can use oxygen, but can survive without it o Strict= obligate= cannot survive with oxygen OR cannot survive without oxygen Ecological Relationships o Free-living= do not depend on another organism for survival o o o o Pathogen= depends on another organism for survival An organism or virus that causes disease Symbiont= provides a benefit for the organism The smaller participant in the symbiotic relationship, living in or on the host Free-living heterotrophs= decomposers Biofilms= surface coating colony of one or more species of prokaryotes that engage in metabolic cooperation Lives on surfaces of water, etc. Community Combination of species Waste products used by the community Organism-of-the-week: Species: Nostoc commune o Common name: star jelly, mare’s eggs, witch’s butter, monster boogers Genus: Nostoc Family: Nostocaceae Order: Nostocales Class: Cyanophyceae Phylum: Cyanobacteria Kingdom: Eubacteria Domain: Bacteria Characteristics: o Cosmopolitan= worldwide o N-fixation= the conversion of atmospheric nitrogen (N2) to ammonia (NH3) N2NH3 o Desiccation resistant Dormancy o Photosynthesis o colonial Domain Archae 1-4+ Kingdoms Kingdom Archaebacteria o Gram negative Lack peptidoglycans Extremophiles o Love extreme conditions o Thermophiles= love extreme temperatures o Halophiles= love salty conditions Methanogens o Produces methane (CH4) as a waste productfossil fuel o Anaerobic Non-extreme archaea Domain Bacteria 1-200+ kingdoms Kingdom Eubacteria o Contains most of the familiar bacteria o Phylum Cyanobacteria Performs photosynthesis Internal membrane Looks like chloroplasts Serial endosymbiotic hypothesis of Eukaryotic organelle origin A hypothesis for the origin of eukaryotes consisting of a sequence of endosymbiotic events in which mitochondria, chloroplasts, and perhaps other cellular structures were derived from small prokaryotes that had been engulfed by larger cells. Endosymbiosis= a relationship between two species in which one organism lives inside the cell or cells of another organism o Proteobacteria Gram negative largest group Escherischia Coli E. Coli Coliformin colon Gut flora Pathogenic strains o Alphaproteobacteria Purple-green Inner membrane looks like mitochondria (ring-shaped DNA) o Spirochaetes Spirilla shape many pathogens o Gram Positive Groups: Chlamydias all are animal diseases Mycoplasmas Smallest of all bacteria No cell wall Ex: walking pneumonia Domain Eukarya 4 kingdoms True nucleus Sexual reproduction o Usually in addition to asexual Why? Because of environmental factors variety in offspring meiosis (mitosis) 3 basic life cycle patterns Meiosis o Replication + 2 divisions o 2nn (1 of each type) o Vocabulary: Reduction division Haploid (half) 23 for humans (n) Diploid 46 for humans (2n) Homologous pair Maternal Paternal Mitosis o 1 replication + 1 division o 2n2n or nn o Gamete Haploid Will fuse= fertilization=syngamy Produced by mitosis or meiosis o Syngamy Fusion of gametes Plasmogamy n+n n+n Karyogamy n+n 2n o zygote diploid o spores dormancy and dispersal n or 2n asexual or sexual mitosis or meiosis do not fuse 3 patterns: o Zygotic Meiosis Fungi and some protista “Zygote undergoes meiosis” Life cycle: o Gametic Meiosis Animalia and some protista “Gametes produced by meiosis” Life cycle: o Sporic Meiosis Plantae and some protista “spores produced by meiosis” “alternation of generations” A life cycle in which there is both a multicellular diploid form, a sporophyte, and a multicellular haploid form, the gametophyte Life cycle: Evolutionary Trends: o (ancestral)(derived) o Asexual reproductionsexual reproduction o Zygotic meiosisgametic and sporic meiosis o Unicellularmulticellular o Isomorphicheteromorphic Isomorphic= referring to alternating generations in plants and some algae in which the sporophytes and gametophytes look alike, although they differ in chromosome number Heteromorphic= referring to a condition in the life cycle of plants and some algae in which the sporophyte and gametophyte generations differ in morphology. o Isogamyanisogamy (oogamy) o Dominant gametophytesporophyte dominant o Homosporyheterospory Homosporous= a plant species that has a single kind of spore, which typically develops into a bisexual gametophyte Heterosporous= a plant species that has two kinds of spores: microspores, which develop into male gametophytes, and megaspores, which develop into female gametophytes. Kingdom Protista “protozoa and algae” o Protozoa=first animal Autotrophs/heterotrophs or mixotrophic Motile/sessile o Sessile= immobile All 3 sexual life cycles o Zygotic, gametic, and sporic meiosis Polyphyletic o 4 or 5 monophyletic clades Most are unicellular Not plants, animals, or fungi Membrane bound nucleus and other organelles An almost abandoned taxon Phylum diplomonadida o Synapamorphic o Double nuclei in equal sizes o Multiple flagella 8 or more o Giardia intestinalis Intestinal parasite in humans Phylum Parabasala o Connected to basal body of flagella Basal body similar to golgi body in humans o Trichomonas vaginalis STD Mostly parasitic Phylum Kinetoplastida o Kinetoplast large mass of DNA in a mitochondrion o Bodo o Trypanosoma brucei Causes African sleeping sickness Transmitted by Tsetse fly Causes anemia Phylum Euglenozoa o Has flagella and chloroplasts o Animal and plant like o Many are photosynthetic o Euglena Nominate genus Nominate=names after Two flagella Stigma Eye like Absorbs light Spot of pigment Alters movement of flagella Light changes the shape of it Positive phototaxis Means moves toward light Inside flagella: 9+2 flagellar and ciliary axoneme doubles Inside basal body: 9+0 Triplets Undulation Movement of flagella Tip-to-base Anterior flagellum o Moves with flagella in front Chloroplast in Euglena 3 membranes o Peranema Undulates flagellum at the tip Phylum Dinoflagellata o Whirly o 2 flagella 1 is longitudinal 1 is circumferential o Gymnodinium Red pigment In gulf water Autotrophic Toxins Bloom Bloom=photosynthetic population Causes red tide o Noctiluca Sea sparkle Bioluminescent In sea water Phylum Apicomplexa o Apical complex At the tip Allows penetration of host cells o Spore forming parasites o Plasmodium Causes malaria Stages of malaria (zygotic meiosis) Sporozite: n, infective stage Trophozoite: feeding cells in RBC Merozoites: product of schizogony (synchrony) Gametocytes: (micro-, macro-), picked up by mosquito Gametes: (in mosquito now): n, fuse by syngamy Oocyst: zygote(2n) Then undergoes meiosis and starts over at sporozoite o Anopheles Vector= an organism that transmits pathogens from one host to another Phylum Ciliophora o Ciliates= type of protist that moves by cilia o Possesses cilia o Paramecium multimicronucleatum Several micro nuclei o Complex structure: Organelles Pellicle= cell membrane + alveoli Oral groove Where feeding begins Cytostome Cell mouth Cytopharynx Leads food particle Food vacuole Cytoproct Cell anus Exocytosis WEV= water expulsion vesicle Contractile vacuole o Osmoregulatory adaptation Parasites in isotonic fluids do not perform osmosis so there is no need for contractile vacuole Membranelle Example: Vorticella Cirrus Examples: Euplotes Phylum Bacillariophyta o Photosynthetic o Common name: diatoms Photosynthetic protist in the stramenopile clade; diatoms have a unique glass like wall made of silicon dioxide embedded in an organic matrix o Cell wall= frustule Silicious (SiO2) Glass like 2 valves o Centric or pennate o Bloom o Diatomaceous earth Used for water filtering o o o o Sandpaper Toothpaste Asexual and sexual reproduction 2ngametic meiosis Mitosis1 valve conserved in each cell Valve picture: Gets as small as 1/3 in size then sexual reproduction occurs and a zygote is formed Phylum Phaeophyta o Brown algae o Macroalgae o Simple multicellularity o Kelp o Sargassum Air filled bladders that help them float o Sporic meiosis o Thallus= a plant body that is not differentiated into stem and leaves and lacks true roots and a vascular system o Picture: Phylum Rhodophyta o Red algae o Agar o Carageenan in ice cream o Ameboid protozoa Amoebozoan= a protist in a clade that includes many species with lobe-shaped or tube-shaped pseudopodia Amebas= with or without tests Test= in foram protists, a porous shell that consists of a single piece of organic material hardened with calcium carbonate. o Shell like exterior structure Pseudopods used for movement and phagocytosis Picture: Phylum Gymnamoeba o Naked- Without tests Phylum Radiolaria o With tests Silicious o Axopodia Needle-like pseudopods o Lobopodia Opposite of axopodia Not found in radiolaria Most common Phylum Foraminifera o Test is full of chambers o –fera: to bear o Calcareous tests (CaCO2) Chalky appearance o Mermaid’s pennies o Reticulopodia= another type of pseudopod o Forms white cliffs, chalk, pink sand in Bermuda o Ooze amoeba parasitic population Types of Pseudopods Reticulopodia – net like Axopodia Lobopodia Phylum Myxomycota (Mycetoza) o Plasmodial slime molds o Myxo = fungus o Characteristics Plasmodium- growth form Multinucleate mass of protoplasm Detritivores= feed on dead stuff Sporic meiosis Complex life cycle Phylum Chlorophyta o Chloro: green o Common name: green algae o In common with plants o Independent solitary cells: Chlamydomonas o Volvox Colonial o Some thallus o Look the same and shaped the same regardless of haploid or diploid=isomorphic o Example: sea grass Zygotic Meiosis Haploid organisms form gametes by mitosis Isogamy – all the gametes are the same Syngamy results in zygote Zygospores undergoes meiosis in spring Phylum Choanoflagellata: common ancestor with animals o structure and DNA most related to animals Sister group of all multicellular organisms Collar – extensions of cell membrane with microvilli that are finger- like foldings Food vacule = filter feeding Organism-of-the-week: Species: Braseni schreberi o Common name: water shield Genus: Braseni Family: Cabombaceae Order: Nymphaeales Class: Eudicot Phylum: Mognoliophyta Kingdom: Plantae Domain: Eukarya Characteristics: o Hydrophilic o Hydrophobic o Mucilage o Anti-herbivore Prevents getting eaten Kingdom Plantae “the land plants” Sporic meiosis o Kingdom plantae has the most complicated of life cycles Most photosynthesis Cell walls o Cellulose- primary structural component Movement by growth Multicellular o Characteristic that distinguishes plants between protists Bryophytes—Super phylum – the oldest group of plants o 3 phyla o Not far from water o Non-vascular Lack vascular tissue Cannot have true plant organs Lack ability to pump water from source Prevents them from getting big because taller means farther from water o Small o No true organs o Require Moist environment o Flagellated sperm Must have water to swim so hence moist environments o Dominant gametophyte generation Asexual o “amphibians” of plant world Phylum Bryophyta o A phyla of bryophytes o Bry- means moss o –phyta means plant o True mosses and relatives o Sporophytes attached to and are dependent on gametophyte o The sporophyte does not perform photosynthesis o Gametophyte undergoes asexual reproduction o Anisogomy= gametes are not the same Egg and sperm o Sperm gets to the egg by splashing out and swimming from antheridium to archegonium o Water is require for sperm dispersal o Zygote (2n) forms in archegonium Gametophyte sporophyte 1st zygote formation o Gametangium= House the cells that become the gamete o Peristome with teeth like structures o Seta = thread like structure o Bryophyta life cycle: Phylum Hepatophyta o Another phyla of the bryophytes o Same life cycle as Bryophyta except: Antheridiophore Structures that bear antheridia (male) Archegoniophore Structures that bear archegonia(female) Sporangium Spore container Same thing as a capsule in Bryophyta Sporophyte is upside down Elatershygroscopic o Aids in spore dispersal o Change based on humidity o Common name: liverworts Doctrine of signatures: idea that God put his signature in organisms so they would look like what they are used for o Lobed thallus With rhizoids Sheet-like structures Dichotomous branching= how the lobes grow Gemmae in gemma cups = for asexual reproduction Phylum Anthocerophyta o Hornwortsthallus gametophyte Horn-like sporophyte o Plants that look like horns o Meiospores dispersed by dehiscence Dehiscence= splitting open to release contents