Download Adelinia02rev nc US

Adelinia02rev(Montoya&Cohen) -- Boraginaceae
Volume 15
Taxon ed.: Giblin
Tech ed.: A. Buback
Page 1 of 4
17 August 2016
North Central US review comments: Freeman (Kans.)
XX.
2015
ADELINIA
*
J. I. Cohen, Syst. Bot. 40: 617, figs. 3B, 4A.
Adeline’s Hound’s tongue
E
Barbara Montoya
James I. Cohen
Herbs, perennial; hispid to strigose to pilose to sericeous.
Roots small to large taproot.
Stems erect, green.
Leaves
cauline and basal, simple, alternate, petiole present; blade
spatulate to lanceolate to deltate, veins conspicuous, margins
entire, planar.
Inflorescences terminal, scorpioid cymes, tip
coiled; 4--9 flowers per cyme, bracts absent.
Pedicels erect.
Flowers actinomorphic, salverform; calyx lobes 5, free, erect,
equal and not toothed in fruit, without spines; corolla blue to
white, tube present, constricted, fornices 5, inserted above
anthers, 2-lobed trapeziform, throat circular, glabrous, corolla
lobes flared, blue with white at base, equal; stamens attached
above base of corolla, filaments cylindric, anthers not
appressed to style, included in corolla; ovary deeply lobed,
style 1, included, gynobasic, stigmas 2, each semidiscoid or
semispheric.
Fruits nutlets, gynobase pyramidal to ovoid, 1/4
length of longest portion of nutlet; 4 or fewer by abortion,
erect, brown, lanceoloid to orbicular, submedial attachment,
Adelinia02rev(Montoya&Cohen) -- Boraginaceae
Volume 15
Taxon ed.: Giblin
Tech ed.: A. Buback
Page 2 of 4
17 August 2016
adaxial attachment scar broadly triangular, slight keel above
scar, surface dull, glochidiate, glochids grappling-hook shaped.
x = 12.
Species 1: w North America.
Adelinia is a monospecific genus that includes a species
formerly in Cynoglossum.
Although this and other New World
species have nutlets similar to those of species of Cynoglossum,
the New World species that have been included in Cynoglossum are
not members of the same clade as the Old World species;
consequently, the New World species are now included in two
genera, Adelinia and Andersonglossum.
These two genera differ
in leaf, pedicel, and nutlet characteristics, with Adelinia
having petiolate cauline leaves, pedicels that are erect in
fruit, and nutlets with a slight keel above the attachment scar.
In contrast, Andersonglossum bears sessile cauline leaves,
pedicels recurved in fruit, and nutlets that lack a keel.
SELECTED REFERENCES
Kelley, R. B.
Jepson Flora Project (eds.)
2013.
Cynoglossum, in
Jepson eFlora,
http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=21750.
Weigend, M., F. Luebert, F. Selvi, G. Brokamp, and H. H. Hilger.
2013.
Multiple origins for Hound’s tongues (Cynoglossum L.) and
Navel Seeds (Omphalodes Mill.) – The phylogeny of the borage
Adelinia02rev(Montoya&Cohen) -- Boraginaceae
Volume 15
Taxon ed.: Giblin
Tech ed.: A. Buback
family (Boraginaceae s.str.).
Page 3 of 4
17 August 2016
Molec. Phylogenet. Evol. 68: 604-
-618.
1.
Adelinia grande
Bot. 40: 617.
2015
(Douglas ex Lehmann) J. I. Cohen. Syst.
*
Pacific hound’s tongue
E
F
Cynoglossum grande Douglas ex Lehmann, Nov. Stirp. Pug. 2: 25.
1830
Herbs to 1 m.
Stems 1(--3), 5--10 mm wide, hispid.
Leaves:
basal: petiole 10.3--21 x 0.3--0.6 cm; blade spatulate to
lanceolate to deltate, (7--)11.6--17 x (4.4--)6.3--11 cm, base
truncate to rounded to cordate, apex acute to mucronate,
abaxially sericeous, adaxially sparsely sericeous; cauline:
petiole 1.5--18.5 x 0.1--0.5 cm; blade lanceolate to deltate,
3.7--19.4 x 1.2--9.3(--11.7) cm, base truncate to rounded to
cordate, apex acute to apiculate, abaxially sericeous, adaxially
sparsely sericeous.
mm.
Inflorescences 3--20 cm.
Pedicels 0--55
Flowers: sepals linear to lanceolate, 5 x 1--2 mm, apex
acute to acuminate, abaxially densely sericeous, adaxially
glabrous; corolla tube 4--7 x 3--4 mm, lobes elliptic, 4--6 x 3-4 mm, margins entire, apex rounded; filaments attached 3--6 mm
from base of corolla, anthers 1.5--2 x 0.5--1 mm; style 4--8 x
0.3 mm; stigma terminal.
Nutlets 4--8 x 4--8 mm, base truncate
to rounded, apex rounded to acute.
Adelinia02rev(Montoya&Cohen) -- Boraginaceae
Volume 15
Taxon ed.: Giblin
Tech ed.: A. Buback
Flowering Feb--May; fruiting Apr--Jun.
Page 4 of 4
17 August 2016
Open to closed
forests, oak woodlands; 0--1700 m; B.C.; Calif., Oreg., Wash.
Of the two western North American species previously
included in Cynoglossum, Adelinia grande has the broader
geographic distribution than the other species, Andersonglossum
occidentale.
The former ranges from southern British Columbia
to California, while the latter is restricted to California and
Oregon.
The two species also differ in corolla, cauline leaf,
and nutlet characteristics.
The corollas of Adelinia grande are
blue and white, while those of Andersonglossum occidentale are
purple to red.