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EVOLUTION cumulative change in the heritable characteristics of a population Evidence of Evolution fossil record homologous structures ex. pentadactyl limb selective breeding Industrial melanism – ex. peppered moths ex. antibiotic resistance ex. finches on Daphne major vestigial structures molecular biology comparative embryology Fossil Record of Horse Evolution Horse Evolution Fossil Formation Fossils Layers Radioactive decay of C-14 and K-40 half-life of C-14 = 5,730 yrs good for fossils 1,000-100,000 ys Used to date fossils half-life of K-40 = 1.25 billion yrs good for fossils older than 100,000 yrs Homologous Structures ex. the pentadactyl limb Selective Breeding Example: Peppered Moth ADAPTIVE RADIATION Hawaiian honey creepers Darwin’s Finches Finches on Daphne Major, Galapagos Daphne Major is a small island Geospiza fortis feeds on seeds Beak size varies & inherited El Nino (83) brings rain & cold – • variety of seeds La Nina (74-77) brings droughts – • few small soft seeds • more large seeds Example: Antibiotic Resistance http://www.youtube.com/watch?v=zjR6L38yReE&safe=active http://www.youtube.com/watch?v=6jBD8xfbf4Y&safe=active Vestigial Structures Comparative Embryology Biochemical Analysis: DNA & Amino Acid Sequences Trends/Patterns of Selection Original bell curve of population: Directional Selection Example: elephant trunk length Stabilizing Selection Example: human birth weight Disruptive selection Example: rabbit fur color on black & white stones, black, white, grey Frequency-dependent Selection https://cubanosune.wordpress.com/2012/06// Sexual Selection http://artscienceofnature.wordpress.com/category/phylum-mollusca/ DIVERGENT EVOLUTION Polar bear Brown bear Species definition: two organisms that can mate and produce fertile offspring CONVERGENT EVOLUTION shark dolphin Counter shading fusiform shape tuna CONVERGENT EVOLUTION Australian marsupials vs. American placentals Coyote, groundhog, spiny anteater Tasmanian devil (ext), wombat, banded anteater Biogeography: Wallace’s Line Bottleneck Effect http://content.answers.com/main/content/wp/en/thumb/a/a9/150px-Polydactyly_01_Lfoot_AP.jpg Founder Effect SPECIATION: allopatric sympatric Speciation mechanisms Gradualism Puntuated Equilibrium Hardy-Weinberg Equilibrium: Allele frequencies remain stable over time. Assumptions: 1. No immigration or emigration 2. No mutations 3. Large population 4. Random mating 5. No natural selection HW equations: p +q = 1 p2 + 2pq + q2 = 1 Hybrid Vigor (aka heterozygous advantage) WARNING COLORATION aposematic coloring MULLERIAN MIMICRY WASP BATESIAN MIMICRY Monarch Viceroy Now thought to be Mullerian mimicry Coral snake and California king snake “Red touch yellow, kill a fellow. Red touch black, venom lack” BATESIAN MIMICRY CAMOUFLAGE CAMOUFLAGE DISRUPTIVE COLORATION EYESPOTS/STARTLE COEVOLUTION Rufous hummingbird & flower Symbiotic: mutualistic Cleaner fish Clown fish and sea anemone Algae & fungi = lichen Bacteria in root nodules of legume Symbiotic: mutualistic Ants & acacia trees Oxpecker bird and ox Aphids & ants Symbiotic: parasitic Strangler fig tapeworm mistletoe Symbiotic: commensalism Remoras and shark Sponge on scallop Barnacles on whale EVOLUTION PATTERNS Revu Disruptive, Stabilizing, Directional Selection, Sexual, Frequency dependnt Divergent vs Convergent evolution Allopatric vs Sympatric speciation Gradualism vs Punctuated Equilibrium Adaptive Radiation Artificial Selection Mimicry and Camouflage Coevolution and Symbiosis